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As for our belief in the origin of species making any difference in descriptive work, I am sure it is incorrect, for I did all my barnacle work under this point of view. Only I often groaned that I was not allowed simply to decide whether a difference was sufficient to deserve a name.

I am glad to hear about Huxley--a wonderful man.

LETTER 345. TO J.D. HOOKER. Wells Terrace, Ilkley, Otley, Yorkshire, Thursday [before December 9th, 1859].

I have read your discussion (345/1. See "Introductory Essay," page c. Darwin did not receive this work until December 23rd, so that the reference is to proof-sheets.), as usual, with great interest. The points are awfully intricate, almost at present beyond the confines of knowledge. The view which I should have looked at as perhaps most probable (though it hardly differs from yours) is that the whole world during the Secondary ages was inhabited by marsupials, araucarias (Mem.--Fossil wood so common of this nature in South America (345/2.

See Letter 6, Note.)), Banksia, etc.; and that these were supplanted and exterminated in the greater area of the north, but were left alive in the south. Whence these very ancient forms originally proceeded seems a hopeless enquiry.



Your remarks on the pa.s.sage of the northern forms southward, and of the southern forms of no kinds pa.s.sing northward, seem to me grand.

Admirable, also, are your remarks on the struggle of vegetation: I find that I have rather misunderstood you, for I feared I differed from you, which I see is hardly the case at all. I cannot help suspecting that you put rather too much weight to climate in the case of Australia. La Plata seems to present such a.n.a.logous facts, though I suppose the naturalisation of European plants has there taken place on a still larger scale than in Australia...

You will get four copies of my book--one for self, and three for the foreign botanists--in about ten days, or sooner; i.e., as soon as the sheets can be bound in cloth. I hope this will not be too late for your parcels.

When you read my volume, use your pencil and score, so that some time I may have a talk with you on any criticisms.

LETTER 346. TO HUGH FALCONER. Down, December 17th, [1859].

Whilst I think of it, let me tell you that years ago I remember seeing in the Museum of the Geological Society a tooth of hippopotamus from Madagascar: this, on geographical and all other grounds, ought to be looked to. Pray make a note of this fact. (346/1. At a meeting of the Geological Society, May 1st, 1833, a letter was read from Mr. Telfair to Sir Alex. Johnstone, accompanying a specimen of recent conglomerate rock, from the island of Madagascar, containing fragments of a tusk, and part of a molar tooth of a hippopotamus ("Proc. Geol. Soc." 1833, page 479). There is a reference to these remains of hippopotamus in a paper by Mr. R.B. Newton in the "Geol. Mag." Volume X., 1893; and in Dr.

Forsyth Major's memoir on Megaladapis Madagascariensis ("Phil. Trans. R.

Soc." Volume 185, page 30, 1894).

Since this letter was written, several bones belonging to two or possibly three species of hippopotamus have been found in Madagascar.

See Forsyth Major, "On the General Results of a Zoological Expedition to Madagascar in 1894-96" ("Proc. Zool. Soc." 1896, page 971.))

We have returned a week ago from Ilkley, and it has done me some decided good. In London I saw Lyell (the poor man who has "rushed into the bosom of two heresies"--by the way, I saw his celts, and how intensely interesting), and he told me that you were very antagonistic to my views on species. I well knew this would be the case. I must freely confess, the difficulties and objections are terrific; but I cannot believe that a false theory would explain, as it seems to me it does explain, so many cla.s.ses of facts. Do you ever see Wollaston? He and you would agree nicely about my book (346/2. "Origin of Species," 1859.)--ill luck to both of you. If you have anything at all pleasant for me to hear, do write; and if all that you can say is very unpleasant, it will do you good to expectorate. And it is well known that you are very fond of writing letters. Farewell, my good old friend and enemy.

Do make a note about the hippopotamus. If you are such a gentleman as to write, pray tell me how Torquay agrees with your health.

(PLATE: DR. ASA GRAY, 1867.)

LETTER 347. TO ASA GRAY. Down, December 24th [1859].

I have been for ten weeks at Water-cure, and on my return a fortnight ago through London I found a copy of your Memoir, and heartily do I thank you for it. (347/1. "Diagnostic Characters of New Species of Phaenogamous Plants collected in j.a.pan by Charles Wright...with Observations upon the Relations of the j.a.panese Flora to that of North America and of other parts of the Northern Temperate Zone" ("Mem.

American Acad. Arts and Sci." Volume VI., page 377, 1857).) I have not read it, and shall not be able very soon, for I am much overworked, and my stomach has got nearly as bad as ever.

With respect to the discussion on climate, I beg you to believe that I never put myself for a moment in compet.i.tion with Dana; but when one has thought on a subject, one cannot avoid forming some opinion. What I wrote to Hooker I forget, after reading only a few sheets of your Memoir, which I saw would be full of interest to me. Hooker asked me to write to you, but, as I told him, I would not presume to express an opinion to you without careful deliberation. What he wrote I know not: I had previously several years ago seen (by whom I forget) some speculation on warmer period in the U. States subsequent to Glacial period; and I had consulted Lyell, who seemed much to doubt, and Lyell's judgment is really admirably cautious. The arguments advanced in your paper and in your letter seem to me hardly sufficient; not that I should be at all sorry to admit this subsequent and intercalated warmer period--the more changes the merrier, I think. On the other hand, I do not believe that introduction of the Old World forms into New World subsequent to the Glacial period will do for the modified or representative forms in the two Worlds. There has been too much change in comparison with the little change of isolated alpine forms; but you will see this in my book. (347/2. "Origin of Species" (1859), Chapter XI., pages 365 et seq.) I may just make a few remarks why at first sight I do not attach much weight to the argument in your letter about the warmer climate. Firstly, about the level of the land having been lower subsequently to Glacial period, as evidenced by the whole, etc., I doubt whether meteorological knowledge is sufficient for this deduction: turning to the S. hemisphere, it might be argued that a greater extent of water made the temperature lower; and when much of the northern land was lower, it would have been covered by the sea and intermigration between Old and New Worlds would have been checked. Secondly, I doubt whether any inference on nature of climate can be deduced from extinct species of mammals. If the musk-ox and deer of great size of your Barren-Grounds had been known only by fossil bones, who would have ventured to surmise the excessively cold climate they lived under? With respect to food of large animals, if you care about the subject will you turn to my discussion on this subject partly in respect to the Elephas primigenius in my "Journal of Researches" (Murray's Home and Colonial Library), Chapter V., page 85. (347/3. "The firm conviction of the necessity of a vegetation possessing a character of tropical luxuriance to support such large animals, and the impossibility of reconciling this with the proximity of perpetual congelation, was one chief cause of the several theories of sudden revolutions of climate...I am far from supposing that the climate has not changed since the period when these animals lived, which now lie buried in the ice. At present I only wish to show that as far as quant.i.ty of food alone is concerned, the ancient rhinoceroses might have roamed over the steppes of Central Siberia even in their present condition, as well as the living rhinoceroses and elephants over the karoos of Southern Africa" ("Journal of Researches,"

page 89, 1888).) In this country we infer from remains of Elephas primigenius that the climate at the period of its embedment was very severe, as seems countenanced by its woolly covering, by the nature of the deposits with angular fragments, the nature of the co-embedded sh.e.l.ls, and co-existence of the musk-ox. I had formerly gathered from Lyell that the relative position of the Megatherium and Mylodon with respect to the Glacial deposits, had not been well made out; but perhaps it has been so recently. Such are my reasons for not as yet admitting the warmer period subsequent to Glacial epoch; but I daresay I may be quite wrong, and shall not be at all sorry to be proved so.

I shall a.s.suredly read your essay with care, for I have seen as yet only a fragment, and very likely some parts, which I could not formerly clearly understand, will be clear enough.

LETTER 348. TO J.D. HOOKER. Down, [December] 26th, [1859].

I have just read with intense interest as far as page xxvi (348/1. For Darwin's impression of the "Introductory Essay to the Tasmanian Flora"

as a whole, see "Life and Letters," II., page 257.), i.e. to where you treat of the Australian Flora itself; and the latter part I remember thinking most of in the proof-sheets. Either you have altered a good deal, or I did not see all or was purblind, for I have been much more interested with all the first part than I was before,--not that I did not like it at first. All seems to me very clearly written, and I have been baulked at only one sentence. I think, on the whole, I like the geological, or rather palaeontological, discussion best: it seems to me excellent, and admirably cautious. I agree with all that you say as far as my want of special knowledge allows me to judge.

I have no criticisms of any importance, but I should have liked more facts in one or two places, which I shall not ask about. I rather demur to the fairness of your comparison of rising and sinking areas (348/2.

Hooker, op. cit., page xv, paragraph 24. Hooker's view was that sinking islands "contain comparatively fewer species and fewer peculiar generic types than those which are rising." In Darwin's copy of the Essay is written on the margin of page xvi: "I doubt whole case."), as in the Indian Ocean you compare volcanic land with exclusively coral islands, and these latter are very small in area and have very peculiar soil, and during their formation are likely to have been utterly submerged, perhaps many times, and restocked with existing plants. In the Pacific, ignorance of Marianne and Caroline and other chief islands almost prevent comparison (348/3. Gambier Island would be an interesting case.

[Note in original.]); and is it right to include American islands like Juan Fernandez and Galapagos? In such lofty and probably ancient islands as Sandwich and Tahiti it cannot make much difference in the flora whether they have sunk or risen a few thousand feet of late ages.

I wish you could work in your notion of certain parts of the Tropics having kept hot, whilst other parts were cooled; I tried this scheme in my mind, and it seemed to fail. On the whole, I like very much all that I have read of your Introduction, and I cannot doubt that it will have great weight in converting other botanists from the doctrine of immutable creation. What a lot of matter there is in one of your pages!

There are many points I wish much to discuss with you.

How I wish you could work out the Pacific floras: I remember ages ago reading some of your MS. In Paris there must be, I should think, materials from French voyages. But of all places in the world I should like to see a good flora of the Sandwich Islands. (348/4. See Hillebrand, "Flora of the Hawaiian Islands," 1888.) I would subscribe 50 pounds to any collector to go there and work at the islands. Would it not pay for a collector to go there, especially if aided by any subscription? It would be a fair occasion to ask for aid from the Government grant of the Royal Society. I think it is the most isolated group in the world, and the islands themselves well isolated from each other.

LETTER 349. TO ASA GRAY. Down, January 7th [1860].

I have just finished your j.a.pan memoir (349/1. "Diagnostic Characters of New Species of Phaenogamous Plants collected in j.a.pan by Charles Wright.

With observations upon the Relations of the j.a.panese Flora to that of North America, etc.: 1857-59."--"Memoirs of Amer. Acad." VI.), and I must thank you for the extreme interest with which I have read it. It seems to me a most curious case of distribution; and how very well you argue, and put the case from a.n.a.logy on the high probability of single centres of creation. That great man Aga.s.siz, when he comes to reason, seems to me as great in taking a wrong view as he is great in observing and cla.s.sifying. One of the points which has struck me as most remarkable and inexplicable in your memoir is the number of monotypic (or nearly so) genera amongst the representative forms of j.a.pan and N. America. And how very singular the preponderance of identical and representative species in Eastern, compared with Western, America. I have no good map showing how wide the moderately low country is on the west side of the Rocky Mountains; nor, of course, do I know whether the whole of the low western territory has been botanised; but it has occurred to me, looking at such maps as I have, that the eastern area must be larger than the western, which would account to a certain small extent for preponderance on eastern side of the representative species.

Is there any truth in this suspicion? Your memoir sets me marvelling and reflecting. I confess I am not able quite to understand your Geology at pages 447, 448; but you would probably not care to hear my difficulties, and therefore I will not trouble you with them.

I was so grieved to get a letter from Dana at Florence, giving me a very poor (though improved) account of his health.

LETTER 350. TO T.H. HUXLEY. 15, Marine Parade, Eastbourne, November 1st [1860].

Your note has been wonderfully interesting. Your term, "pithecoid man,"

is a whole paper and theory in itself. How I hope the skull of the new Macrauchenia has come. It is grand. I return Hooker's letter, with very many thanks. The glacial action on Lebanon is particularly interesting, considering its position between Europe and Himalaya. I get more and more convinced that my doctrine of mundane Glacial period is correct (350/1. In the 1st edition of the "Origin," page 373, Darwin argues in favour of a Glacial period practically simultaneous over the globe. In the 5th edition, 1869, page 451, he adopted Mr. Croll's views on the alternation of cold periods in the northern and southern hemispheres. An interesting modification of the mundane Glacial period theory is given in Belt's "The Naturalist in Nicaragua," 1874, page 265. Mr. Belt's views are discussed in Wallace's "Geogr. Distribution," 1876, Volume I., page 151.), and that it is the most important of all late phenomena with respect to distribution of plants and animals. I hope your Review (350/2. The history of the foundation of the "Natural History Review"

is given in Huxley's "Life and Letters," Volume I., page 209. See Letter 107.) progresses favourably. I am exhausted and not well, so write briefly; for we have had nine days of as much misery as man can endure.

My poor daughter has suffered pitiably, and night and day required three persons to support her. The crisis of extreme danger is over, and she is rallying surprisingly, but the doctors are yet doubtful of ultimate issue. But the suffering was so pitiable I almost got to wish to see her die. She is easy now. When she will be fit to travel home I know not.

I most sincerely hope that Mrs. Huxley keeps up pretty well. The work which most men have to do is a blessing to them in such cases as yours.

G.o.d bless you.

Sir H. Holland came here to see her, and was wonderfully kind.

LETTER 351. TO C. LYELL. Down, November 20th [1860].

I quite agree in admiration of Forbes' Essay (351/1. "Memoir of the Geolog. Survey of the United Kingdom," Volume I., 1846.), yet, on my life, I think it has done, in some respects, as much mischief as good.

Those who believe in vast continental extensions will never investigate means of distribution. Good heavens, look at Heer's map of Atlantis!

I thought his division and lines of travel of the British plants very wild, and with hardly any foundation. I quite agree with what you say of almost certainty of Glacial epoch having destroyed the Spanish saxifrages, etc., in Ireland. (351/2. See Letter 20.) I remember well discussing this with Hooker; and I suggested that a slightly different or more equable and humid climate might have allowed (with perhaps some extension of land) the plants in question to have grown along the entire western sh.o.r.es between Spain and Ireland, and that subsequently they became extinct, except at the present points under an oceanic climate.

The point of Devonshire now has a touch of the same character.

I demur in this particular case to Forbes' transportal by ice. The subject has rather gone out of my mind, and it is not worth looking to my MS. discussion on migration during the Glacial period; but I remember that the distribution of mammalia, and the very regular relation of the Alpine plants to points due north (alluded to in "Origin"), seemed to indicate continuous land at close of Glacial period.

LETTER 352. TO J.D. HOOKER. Down, March 18th [1861].

I have been recalling my thoughts on the question whether the Glacial period affected the whole world contemporaneously, or only one longitudinal belt after another. To my sorrow my old reasons for rejecting the latter alternative seem to me sufficient, and I should very much like to know what you think. Let us suppose that the cold affected the two Americas either before or after the Old World. Let it advance first either from north or south till the Tropics became slightly cooled, and a few temperate forms reached the Silla of Caracas and the mountains of Brazil. You would say, I suppose, that nearly all the tropical productions would be killed; and that subsequently, after the cold had moderated, tropical plants immigrated from the other non-chilled parts of the world. But this is impossible unless you bridge over the tropical parts of the Atlantic--a doctrine which you know I cannot admit, though in some respects wishing I could. Oswald Heer would make nothing of such a bridge. When the Glacial period affected the Old World, would it not be rather rash to suppose that the meridian of India, the Malay Archipelago, and Australia were refrigerated, and Africa not refrigerated? But let us grant that this was so; let us bridge over the Red Sea (though rather opposed to the former almost certain communication between the Red Sea and the Mediterranean); let us grant that Arabia and Persia were damp and fit for the pa.s.sage of tropical plants: nevertheless, just look at the globe and fancy the cold slowly coming on, and the plants under the tropics travelling towards the equator, and it seems to me highly improbable that they could escape from India to the still hot regions of Africa, for they would have to go westward with a little northing round the northern sh.o.r.es of the Indian Ocean. So if Africa were refrigerated first, there would be considerable difficulty in the tropical productions of Africa escaping into the still hot regions of India. Here again you would have to bridge over the Indian Ocean within so very recent a period, and not in the line of the Laccadive Archipelago. If you suppose the cold to travel from the southern pole northwards, it will not help us, unless we suppose that the countries immediately north of the northern tropic were at the same time warmer, so as to allow free pa.s.sage from India to Africa, which seems to me too complex and unsupported an hypothesis to admit.

Therefore I cannot see that the supposition of different longitudinal belts of the world being cooled at different periods helps us much.

The supposition of the whole world being cooled contemporaneously (but perhaps not quite equally, South America being less cooled than the Old World) seems to me the simplest hypothesis, and does not add to the great difficulty of all the tropical productions not having been exterminated. I still think that a few species of each still existing tropical genus must have survived in the hottest or most favourable spots, either dry or damp. The tropical productions, though much distressed by the fall of temperature, would still be under the same conditions of the length of the day, etc., and would be still exposed to nearly the same enemies, as insects and other animals; whereas the invading temperate productions, though finding a favouring temperature, would have some of their conditions of life new, and would be exposed to many new enemies. But I fully admit the difficulty to be very great.

I cannot see the full force of your difficulty of no known cause of a mundane change of temperature. We know no cause of continental elevations and depressions, yet we admit them. Can you believe, looking to Europe alone, that the intense cold, which must have prevailed when such gigantic glaciers extended on the plains of N. Italy, was due merely to changed positions of land within so recent a period? I cannot.

It would be far too long a story, but it could, I think, be clearly shown that all our continents existed approximately in their present positions long before the Glacial period; which seems opposed to such gigantic geographical changes necessary to cause such a vast fall of temperature. The Glacial period endured in Europe and North America whilst the level of the land oscillated in height fully 3,000 feet, and this does not look as if changed level was the cause of the Glacial period. But I have written an unreasonably long discussion. Do not answer me at length, but send me a few words some time on the subject.

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