Little Masterpieces of Science Part 1

Little Masterpieces of Science.

by Various.

PREFACE

To gather stones and fallen boughs is soon to ask, what may be done with them, can they be piled and fastened together for shelter? So begins architecture, with the hut as its first step, with the Alhambra, St.

Peter's, the capitol at Washington, as its last. In like fashion the ama.s.sing of fact suggests the ordering of fact: when observation is sufficiently full and varied it comes to the reasons for what it sees.

The geologist delves from layer to layer of the earth beneath his tread, he finds as he compares their fossils that the more recent forms of life stand highest in the scale of being, that in the main the animals and plants of one era are more allied to those immediately next than to those of remoter times. He thus divines that he is but exploring the proofs of lineal descent, and with this thought in his mind he finds that the collections not only of his own district, but of every other, take on a new meaning. The great seers of science do not await every jot and t.i.ttle of evidence in such a case as this. They discern the drift of a fact here, a disclosure there, and with both wisdom and boldness a.s.sume that what they see is but a promise of what shall duly be revealed. Thus it was that Darwin early in his studies became convinced of the truth of organic evolution: the labours of a lifetime of all but superhuman effort, a judicial faculty never exceeded among men, served only to confirm his confidence that all the varied forms of life upon earth have come to be what they are through an intelligible process, mainly by "natural selection."

The present volume offers from the cla.s.sic pages of Darwin his summary of the argument of "The Origin of Species," his account of how that book came to be written, and his recapitulation of "The Descent of Man." All this affords a supreme lesson as to the value of observation with a purpose. When Darwin was confronted with an organ or trait which puzzled him, he was wont to ask, What use can it have had? And always the answer was that every new peculiarity of plant, or beast, is seized upon and held whenever it confers advantage in the unceasing conflict for place and food. No hue of scale or plume, no curve of beak or note of song, but has served a purpose in the plot of life, or advanced the action in a drama where the penalty for failure is extinction.

As Charles Darwin stood first among the naturalists of the nineteenth century, his advocacy of evolution soon wrought conviction among the thinkers competent to follow his evidence and weigh his arguments. The opposition to his theories though short was sharp, and here he found a lieutenant of unflinching courage, of the highest expository power, in Professor Huxley. This great teacher came to America in 1876, and discoursed on the ancestry of the horse, as disclosed in fossils then recently discovered in the Far West, maintaining that they afforded unimpeachable proof of organic evolution. His princ.i.p.al lecture is here given.

In a remarkable field of "natural selection" Bates, Wallace and Poulton have explained the value of "mimicry" as an aid to beasts, birds, insects, as they elude their enemies or lie unsuspected on the watch for prey. The resemblances thus worked out through successive generations attest the astonishing plasticity of bodily forms, a plasticity which would be incredible were not its evidence under our eyes in every quarter of the globe. Insects have high economic importance as agents of destruction: we are learning how to pit one of them against another, so as to leave a clear field to the farmer and the fruit grower. In this department a leader is Professor Howard, who contributes a noteworthy chapter on the successful fight against the pest which threatened with ruin the orange groves of California.

To the every-day observer the most enticing field of natural history is that in which common flowers and common insects work out their unending co-partnery. A blossom by its scent, its beauty of tint, allures a moth or bee and thus, in effect, is able to take flight and find a mate across a county so as to perpetuate its race a hundred miles from home.

Our volume closes with a sketch of the singular ties which thus bind together the fortunes of blossom and insect, so that at last the very form of a flower may be cast in the mould of its winged ally. A word is also spoken regarding the singular relations of late detected between the world of vegetation and minute forms once deemed parasitic. The pea and its kindred harbor on their rootlets certain tiny lodgers; the tenants pay a liberal rent in the form of nitrogen compounds, a striking interlacement of interests!

GEORGE ILES.

THE NATURALIST AS INTERPRETER AND SEER

THE ORIGIN OF SPECIES: THE ARGUMENT IN SUMMARY

CHARLES DARWIN

[Charles Darwin, one of the greatest men of all time, did more to advance and prove the theory of evolution than anybody else who ever lived. This he accomplished by virtue of the highest gifts of observation, experiment, and generalization. His truthfulness, patience, and calmness of judgment have never been exceeded by mortal. His works are published by D. Appleton & Co., New York, together with his "Life and Letters," edited by his son Francis. From "The Origin of Species" the argument in summary is here given.]

On the view that species are only strongly marked and permanent varieties, and that each species first existed as a variety, we can see why it is that no line of demarcation can be drawn between species, commonly supposed to have been produced by special acts of creation, and varieties which are acknowledged to have been produced by secondary laws. On this same view we can understand how it is that in a region where many species of a genus have been produced, and where they now flourish, these same species should present many varieties; for where the manufactory of species has been active, we might expect, as a general rule, to find it still in action; and this is the case if varieties be incipient species. Moreover, the species of the larger genera, which afford the greater number of varieties or incipient species, retain to a certain degree the character of varieties; for they differ from each other by a less amount of difference than do the species of smaller genera. The closely allied species also of a larger genera apparently have restricted ranges, and in their affinities they are cl.u.s.tered in little groups round other species--in both respects resembling varieties. These are strange relations on the view that each species was independently created, but are intelligible if each existed first as a variety.

As each species tends by its geometrical rate of reproduction to increase inordinately in number; and as the modified descendants of each species will be enabled to increase by as much as they become more diversified in habits and structure, so as to be able to seize on many and widely different places in the economy of nature, there will be a constant tendency in natural selection to preserve the most divergent offspring of any one species. Hence, during a long-continued course of modification, the slight differences of characteristic of varieties of the same species, tend to be augmented into the greater differences characteristic of the species of the same genus. New and improved varieties will inevitably supplant and exterminate the older, less improved, and intermediate varieties; and thus species are rendered to a large extent defined and distinct objects. Dominant species belonging to the larger groups within each cla.s.s tend to give birth to new and dominant forms; so that each large group tends to become still larger, and at the same time more divergent in character. But as all groups cannot thus go on increasing in size, for the world would not hold them, the more dominant groups beat the less dominant. This tendency in the large groups to go on increasing in size and diverging in character, together with the inevitable contingency of much extinction, explains the arrangement of all the forms of life in groups subordinate to groups, all within a few great cla.s.ses, which has prevailed throughout all time. This grand fact of the grouping of all organic beings under what is called the Natural System, is utterly inexplicable on the theory of creation.

As natural selection acts solely by acc.u.mulating slight, successive, favourable variations, it can produce no great or sudden modifications; it can act only by short and slow steps. Hence, the canon of "Nature makes no leaps," which every fresh addition to our knowledge tends to confirm, is on this theory intelligible. We can see why throughout nature the same general end is gained by an almost infinite diversity of means, for every peculiarity when once acquired is long inherited, and structures already modified in many different ways have to be adapted for the same general purpose. We can, in short, see why nature is prodigal in variety, though n.i.g.g.ard in innovation. But why this should be a law of nature if each species has been independently created no man can explain.

Many other facts are, as it seems to me, explicable on this theory. How strange it is that a bird, under the form of a woodp.e.c.k.e.r, should prey on insects on the ground; that upland geese which rarely or never swim, would possess webbed feet; that a thrush-like bird should dive and feed on sub-aquatic insects; and that a petrel should have the habits and structure fitting it for the life of an auk! and so in endless other cases. But on the view of each species constantly trying to increase in number, with natural selection always ready to adapt the slowly varying descendants of each to any unoccupied or ill-occupied place in nature, these facts cease to be strange, or might even have been antic.i.p.ated.

We can to a certain extent understand how it is that there is so much beauty throughout nature; for this may be largely attributed to the agency of selection. That beauty, according to our sense of it, is not universal, must be admitted by every one who will look at some venomous snakes, at some fishes, and at certain hideous bats with a distorted resemblance to the human face. s.e.xual selection has given the most brilliant colours, elegant patterns, and other ornaments to the males, and sometimes to both s.e.xes of many birds, b.u.t.terflies and other animals. With birds it has often rendered the voice of the male musical to the female, as well as to our ears. Flowers and fruit have been rendered conspicuous by brilliant colours in contrast with the green foliage, in order that the flowers may be easily seen, visited and fertilized by insects, and the seeds disseminated by birds. How it comes that certain colours, sounds and forms should give pleasure to man and the lower animals, that is, how the sense of beauty in its simplest form was first acquired, we do not know any more than how certain odours and flavours were first rendered agreeable.

As natural selection acts by compet.i.tion, it adopts and improves the inhabitants of each country only in relation to their co-inhabitants; so that we need feel no surprise at the species of any one country, although on the ordinary view supposed to have been created and specially adapted for that country, being beaten and supplanted by the naturalized productions from another land. Nor ought we marvel if all the contrivances in nature be not, as far as we can judge, absolutely perfect, as in the case even of the human eye; or if some of them be abhorrent to our ideas of fitness. We need not marvel at the sting of the bee, when used against an enemy, causing the bee's own death; at drones being produced in such great numbers for one single act, and being then slaughtered by their sterile sisters; at the astonishing waste of pollen by our fir trees; at the instinctive hatred of the queen bee for her own fertile daughters; at ichneumonidae feeding within the living bodies of caterpillars; or at other such cases. The wonder indeed, is, on the theory of natural selection, that more cases of the want of absolute perfection have not been detected.

The complex and little known laws governing production of varieties are the same, as far as we can judge, with the laws which have governed the production of distinct species. In both cases physical conditions seem to have produced some direct and definite effect, but how much we cannot say. Thus, when varieties enter any new station, they occasionally a.s.sume some of the characters proper to the species of that station.

With both varieties and species, use and disuse seem to have produced a considerable effect; for it is impossible to resist this conclusion when we look, for instance, at the logger-headed duck, which has wings incapable of flight, in nearly the same condition as in the domestic duck; or when we look at the burrowing tucu-tucu, which is occasionally blind, and then at certain moles, which are habitually blind and have their eyes covered with skin; or when we look at the blind animals inhabiting the dark caves of America and Europe. With varieties and species, correlated variation seems to have played an important part, so that when one part has been modified other parts have been necessarily modified. With both varieties and species, reversions to long-lost characters occasionally occur. How inexplicable on the theory of creation is the occasional appearance of stripes on the shoulders and legs of the several species of the horse-genus and of their hybrids! How simply is this fact explained if we believe that these species are all descended from a striped progenitor, in the same manner as the several domestic breeds of the pigeon are descended from the blue and barred rock pigeon!

On the ordinary view of each species having been independently created, why should specific characters, or those by which the species of the same genus differ from each other, be more variable than generic characters in which they all agree? Why, for instance, should the colour of a flower be more likely to vary in any one species of genus, if the other species possess differently coloured flowers, than if all possessed the same coloured flowers? If species are only well-marked varieties, of which the characters have become in a high degree permanent, we can understand this fact; for they have already varied since they branched off from a common progenitor in certain characters, by which they have come to be specifically different from each other; therefore these same characters would be more likely again to vary than the generic characters which have been inherited without change for an immense period. It is inexplicable on the theory of creation why a part developed in a very unusual manner in one species alone of a genus, and therefore, as we may naturally infer, of great importance to that species, should be eminently liable to variation; but, on our view, this part has undergone, since the several species branched off from a common progenitor, an unusual amount of variability and modification, and therefore we might expect the part generally to be still variable.

But a part may be developed in the most unusual manner, like the wing of a bat, and yet not be more variable than any other structure, if the part be common to many subordinate forms, that is, if it has been inherited for a very long period; for in this case it will have been rendered constant by long-continued natural selection.

Glancing at instincts, marvellous as some are, they offer no greater difficulty than do corporeal structures on the theory of the natural selection of successive, slight, but profitable modifications. We can thus understand why nature moves by graduated steps in endowing certain animals of the same cla.s.s with their several instincts. I have attempted to show how much light the principle of gradation throws on the admirable architectural powers of the hive-bee. Habit no doubt often comes into play in modifying instincts; but it certainly is not indispensable, as we see in the case of neuter insects, which leave no progeny to inherit the effects of long-continued habit. On the view of all the species of the same genus having descended from a common parent, and having inherited much in common, we can understand how it is that allied species, when placed under widely different conditions of life, yet follow nearly the same instincts; why the thrushes of temperate and tropical South America, for instance, line their nests with mud like our British species. On the view of instincts having been slowly acquired through natural selection, we need not marvel at some instincts being not perfect and liable to mistakes, and at many instincts causing other animals to suffer.

If species be only well-marked and permanent varieties, we can see at once why their crossed offspring should follow the same complex laws in their degrees and kinds of resemblance to their parents--in being absorbed into each other by successive crosses, and in other such points--as do the crossed offspring of acknowledged varieties. This similarity would be a strange fact, if species had been independently created and varieties had been produced through secondary laws.

If we admit that the geological record is imperfect to an extreme degree, then the facts, which the record does give, strongly support the theory of descent with modification. New species have come on the stage slowly and at successive intervals; and the amount of change after equal intervals of time, is widely different in different groups. The extinction of species and of whole groups of species, which has played so conspicuous a part in the history of the organic world, almost inevitably follows from the principle of natural selection; for old forms are supplanted by new and improved forms. Neither single species nor groups of species reappear when the chain of ordinary generation is once broken. The gradual diffusion of dominant forms, with the slow modification of their descendants, causes the forms of life, after long intervals of time, to appear as if they had changed simultaneously throughout the world. The fact of the fossil remains of each formation being in some degree intermediate in character between the fossils in the formations above and below, is simply explained by their intermediate position in the chain of descent. The grand fact that all extinct beings can be cla.s.sed with all recent beings, naturally follows from the living and the extinct being the offspring of common parents.

As species have generally diverged in character during their long course of descent and modification, we can understand why it is that the more ancient forms, or early progenitors of each group, so often occupy a position in some degree intermediate between existing groups. Recent forms are generally looked upon as being, on the whole, higher in the scale of organization than ancient forms; and they must be higher, in so far as the later and more improved forms have conquered the older and less improved forms in the struggle for life; they have also generally had their organs more specialized for different functions. This fact is perfectly compatible with numerous beings still retaining simple but little improved structures, fitted for simple conditions of life; it is likewise compatible with some forms having retrograded in organization, by having become at each stage of descent better fitted for new and degraded habits of life. Lastly, the wonderful law of the long endurance of allied forms on the same continent--of marsupials [as kangaroos] in Australia, of edentata [as armadillos, sloths, and anteaters] in America, and other such cases--is intelligible, for within the same country the existing and the extinct will be closely allied by descent.

Looking to geographical distribution, if we admit that there has been during the long course of ages much migration from one part of the world to another, owing to former climatical and geographical changes and to the many occasional and unknown means of dispersal, then we can understand, on the theory of descent with modification, most of the great leading facts in distribution. We can see why there should be so striking a parallelism in the distribution of organic beings throughout s.p.a.ce, and in their geological succession throughout time; for in both cases the beings have been connected by the bond of ordinary generation, and the means of modification have been the same. We see the full meaning of the wonderful fact, which has struck every traveller, namely, that on the same continent, under the most diverse conditions, under heat and cold, on mountain and lowland, on deserts and marshes, most of the inhabitants within each great cla.s.s are plainly related; for they are the descendants of the same progenitors and early colonists. On this same principle of former migration, combined in most cases with modification, we can understand by the aid of the Glacial period, the ident.i.ty of some few plants and the close alliance of many others, on the most distant mountains, and in the northern and southern temperate zones; and likewise the close alliance of some of the inhabitants of the sea in the northern and southern temperate lat.i.tudes, though separated by the whole inter-tropical ocean. Although two countries may present physical conditions as closely similar as the same species ever acquire, we need feel no surprise at their inhabitants being widely different, if they have been for a long period completely sundered from each other; for as the relation of organism to organism is the most important of all relations, and as the two countries will have received colonists at various periods and in different proportions, from some other country or from each other, the course of modification in the two areas will inevitably have been different.

On this view of migration, with subsequent modification, we see why oceanic islands are inhabited by only few species, but of these, why many are peculiar or endemic forms. We clearly see why species belonging to those groups of animals which cannot cross wide s.p.a.ces of the ocean, as frogs and terrestrial mammals, do not inhabit oceanic islands; and why, on the other hand, new and peculiar species of bats, animals which can traverse the ocean, are often found on islands far distant from any continent. Such cases as the presence of peculiar species of bats on oceanic islands and the absence of all other terrestrial mammals, are facts utterly inexplicable on the theory of independent acts of creation.

The existence of closely allied representative species in any two areas, implies on the theory of descent with modification, that the same parent-forms formerly inhabited both areas: and we almost invariably find that wherever many closely allied species inhabit two areas, some identical species are still common to both. Wherever many closely allied yet distant species occur, doubtful forms and varieties belonging to the same groups likewise occur. It is a rule of high generality that the inhabitants of each area are related to the inhabitants of the nearest source whence immigrants might have been derived. We see this in the striking relation of nearly all the plants and animals of the Galapagos Archipelago, of Juan Fernandez, and of the other American islands, to the plants and animals of the neighbouring American mainland; and of those of the Cape Verde Archipelago, and of the other African islands to the African mainland. It must be admitted that these facts receive no explanation on the theory of creation.

The fact, as we have seen, that all past and present organic beings can be arranged within a few great cla.s.ses, in groups subordinate to groups, and with the extinct groups often falling in between the recent groups, is intelligible on the theory of natural selection with its contingencies of extinction and divergence of character. On these same principles we see how it is that the mutual affinities of the forms within each cla.s.s are so complex and circuitous. We see why certain characters are far more serviceable than others for cla.s.sification; why adaptive characters derived from rudimentary parts, though of no service to the beings, are often of high cla.s.sificatory value; and why embryological characters are often the most valuable of all. The real affinities of all organic beings, in contradistinction to their adaptive resemblances, are due to inheritance or community of descent. The Natural System is a genealogical arrangement, with the acquired grades of difference, marked by the terms, varieties, species, genera, families, etc.; and we have to discover the lines of descent by the most permanent characters, whatever they may be, and of however slight vital importance.

The similar framework of bones in the hand of a man, wing of a bat, fin of the porpoise, and leg of the horse--the same number of vertebrae forming the neck of the giraffe and of the elephant--and innumerable other such facts, at once explain themselves on the theory of descent with slow and slight successive modifications. The similarity of pattern in the wing and in the leg of a bat, though used for such different purpose--in the jaws and legs of a crab--in the petals, stamens, and pistils of a flower, is likewise, to a large extent, intelligible on the view of the gradual modification of parts or organs, which were aboriginally alike in an early progenitor in each of these cla.s.ses. On the principle of successive variations not always supervening at an early age, and being inherited at a corresponding not early period of life, we clearly see why the embryos of mammals, birds, reptiles, and fishes should be so closely similar, and so unlike the adult forms. We may cease marvelling at the embryo of an air-breathing mammal or bird having branchial slits and arteries running in loops, like those of a fish which has to breathe the air dissolved in water by the aid of well-developed branchiae [gills].

Disuse, aided sometimes by natural selection, will often have reduced organs when rendered useless under changed habits or conditions of life; and we can understand on this view the meaning of rudimentary organs.

But disuse and selection will generally act on each creature, when it has come to maturity and has to play its full part in the struggle for existence, and will thus have little power in an organ during early life; hence the organ will not be reduced or rendered rudimentary at this early age. The calf, for instance, has inherited teeth, which never cut through the gums of the upper jaw, from an early progenitor having well-developed teeth; and we may believe, that the teeth in the mature animal were formerly reduced by disuse, owing to the tongue and palate, or lips, having become excellently fitted through natural selection to browse without their aid; whereas in the calf, the teeth have been left unaffected, and on the principle of inheritance at corresponding ages have been inherited from a remote period to the present day. On the view of each organism with all its separate parts having been specially created, how utterly inexplicable is it that organs bearing the plain stamp of inutility, such as the teeth in the embryonic calf or the shrivelled wings under the soldered wing covers of many beetles, should so frequently occur. Nature may be said to have taken pains to reveal her scheme of modification, by means of rudimentary organs, of embryological and h.o.m.ologous [corresponding] structures, but we are too blind to understand her meaning.

I have now recapitulated the facts and considerations which have thoroughly convinced me that species have been modified, during a long course of descent. This has been effected chiefly through the natural selection of numerous successive, slight, favourable variations; aided in an important manner by the inherited effects of the use and disuse of parts; and in an unimportant manner, that is, in relation to adaptive structures, whether past or present, by the direct action of external conditions, and by variations which seem to us in our ignorance to arise spontaneously. It appears that I formerly underrated the frequency and value of these latter forms of variation, as leading to permanent modifications of structure independently of natural selection. But as my conclusions have lately been much misrepresented, and it has been stated that I attribute the modification of species exclusively to natural selection, I may be permitted to remark that in the first edition of this work, and subsequently, I placed in a most conspicuous, position--namely, at the close of the Introduction--the following words: "I am convinced that natural selection has been the main but not the exclusive means of modification." This has been of no avail. Great is the power of steady misrepresentation; but the history of science shows that fortunately this power does not long endure.

It can hardly be supposed that a false theory would explain, in so satisfactory a manner as does the theory of natural selection, the several large cla.s.ses of facts above specified. It has recently been objected that this is an unsafe method of arguing; but it is a method used in judging the common events of life, and has often been used by the greatest natural philosophers. The undulatory theory of light has thus been arrived at; and the belief in the revolution of the earth on its own axis was until lately supported by hardly any direct evidence.

It is no valid objection that science as yet throws no light on the far higher problems of the essence of the origin of life. Who can explain what is the essence of the attraction of gravity? No one now objects to following out the results consequent on this unknown element of attraction; notwithstanding that Leibnitz formerly accused Newton of introducing "occult qualities and miracles into philosophy."

I see no good reasons why the views given in this volume should shock the religious feelings of any one. It is satisfactory, as showing how transient such impressions are, to remember that the greatest discovery ever made by man, namely, the law of the attraction of gravity, was also attacked by Leibnitz, "as subversive of natural, and inferentially of revealed religion." A celebrated author and divine has written to me that "he has gradually learned to see that it is just as n.o.ble a conception of the Deity to believe that He created a few original forms capable of self-development into other and needful forms, as to believe that He required a fresh act of creation to supply the voids caused by the action of His laws."

Why, it may be asked, until recently did nearly all the most eminent living naturalists and geologists disbelieve in the mutability of species? It cannot be a.s.serted that organic beings in a state of nature are subject to no variation; it cannot be proved that the amount of variation in the course of long ages is a limited quant.i.ty; no clear distinction has been, or can be, drawn between species and well-marked varieties. It cannot be maintained that species when intercrossed are invariably sterile and varieties invariably fertile; or that sterility is a special endowment and sign of creation. The belief that species were immutable productions was almost unavoidable as long as the history of the world was thought to be of short duration; and now that we have acquired some idea of the lapse of time, we are too apt to a.s.sume, without proof, that the geological record is so perfect that it would have afforded us plain evidence of the mutation of species, if they had undergone mutation.

But the chief cause of our natural unwillingness to admit that one species has given birth to other and distinct species, is that we are always slow in admitting great changes of which we do not see the steps.

The difficulty is the same as that felt by so many geologists, when Lyell first insisted that long lines of inland cliffs had been formed, and great valleys excavated, by the agencies which we still see at work.

The mind cannot possibly grasp the full meaning of the term of even a million years; it cannot add up and perceive the full effects of many slight variations, acc.u.mulated during an almost infinite number of generations.

Although I am fully convinced of the truth of the views given in this volume under the form of an abstract, I by no means expect to convince experienced naturalists whose minds are stocked with a mult.i.tude of facts all viewed, during a long course of years, from a point of view directly opposite to mine. It is so easy to hide our ignorance under such expressions as the "plan of creation," "unity of design," etc., and to think that we give an explanation when we only restate a fact. Any one whose disposition leads him to attach more weight to unexplained difficulties than to the explanation of a certain number of facts will certainly reject the theory. A few naturalists, endowed with much flexibility of mind, and who have already begun to doubt the immutability of species, may be influenced by this volume; but I look with confidence to the future, to young and rising naturalists, who will be able to view both sides of the question with impartiality. Whoever is led to believe that species are mutable will do good service by conscientiously expressing his conviction; for thus only can the load of prejudice by which this subject is overwhelmed be removed.

Several eminent naturalists have of late published their belief that a mult.i.tude of reputed species in each genus are not real species; but that other species are real, that is, have been independently created.

This seems to me a strange conclusion to arrive at. They admit that a mult.i.tude of forms, which till lately they themselves thought were special creations, and which are still thus looked at by the majority of naturalists, and which consequently have all the external characteristic features of true species--they admit that these have been produced by variation, but they refuse to extend the same view to other and slightly different forms. Nevertheless, they do not pretend that they can define, or even conjecture, which are the created forms of life, and which are those produced by secondary laws. They admit variation as a true cause in one case, they arbitrarily reject it in another, without a.s.signing any distinction in the two cases. The day will come when this will be given as a curious ill.u.s.tration of the blindness of preconceived opinion. These authors seem no more startled at a miraculous act of creation than at an ordinary birth. But do they really believe that at innumerable periods in the earth's history certain elemental atoms have been commanded suddenly to flash into living tissues? Do they believe that at each supposed act of creation one individual or many were produced? Were all the infinite numerous kinds of animals and plants created as eggs or seed, or as full grown? and in the case of mammals, were they created bearing the false marks of nourishment from the mother's womb? Undoubtedly some of these same questions cannot be answered by those who believe in the appearance or creation of only a few forms of life, or of some one form alone. It has been maintained by several authors that it is as easy to believe in the creation of a million beings as of one; but Maupertuis's philosophical axiom "of least action" leads the mind more willingly to admit the smaller number; and certainly we ought not to believe that innumerable beings within each great cla.s.s have been created with plain, but deceptive, marks of descent from a single parent.

As a record of a former state of things, I have retained in the foregoing paragraphs, and elsewhere, several sentences which imply that naturalists believe in the separate creation of each species; and I have been much censured for having thus expressed myself. But undoubtedly this was the general belief when the first edition of the present work appeared. I formerly spoke to very many naturalists on the subject of evolution, and never once met with any sympathetic agreement. It is probable that some did then believe in evolution, but they were either silent or expressed themselves so ambiguously that it was not easy to understand their meaning. Now, things are wholly changed, and almost every naturalist admits the great principle of evolution. There are, however, some who still think that species have suddenly given birth, through quite unexplained means, to new and totally different forms.

But, as I have attempted to show, weighty evidence can be opposed to the admission of great and abrupt modifications. Under a scientific point of view, and as leading to further investigation, but little advantage is gained by believing that new forms are suddenly developed in an inexplicable manner from old and widely different forms, over the old belief in the creation of species from the dust of the earth.

It may be asked how far I extend the doctrine of the modification of species. The question is difficult to answer, because the more distinct the forms are which we consider, by so much the arguments in favour of community of descent become fewer in number and less in force. But some arguments of the greatest weight extend very far. All the members of whole cla.s.ses are connected together by a chain of affinities, and all can be cla.s.sed on the same principle, in groups subordinate to groups.

Fossil remains sometimes tend to fill up very wide intervals between existing orders.