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Miohippus and Anchitherium, from the miocene, were about as large as a sheep. Hipparion and Pliohippus, of the pliocene, equalled the a.s.s in height; while the size of the quaternary Equus was fully up to that of a modern horse.
"The increase of speed was equally well marked, and was a direct result of the gradual formation of the limbs. The latter were slowly concentrated by the reduction of their lateral elements and enlargement of the axial bone, until the force exerted by each limb came to act directly through its axis in the line of motion. This concentration is well seen--e.g., in the fore-limb. There was, first, a change in the scapula and humerus, especially in the latter, which facilitated motion in one line only; second, an expansion of the radius and reduction of the ulna, until the former alone remained entire and effective; third, a shortening of all the carpal bones and enlargement of the median ones, insuring a firmer wrist; fourth, an increase of size of the third digit, at the expense of those of each side, until the former alone supported the limb.
"Such is, in brief, a general outline of the more marked changes that seemed to have produced in America the highly specialized modern Equus from his diminutive four-toed predecessor, the eocene Orohippus. The line of descent appears to have been direct, and the remains now known supply every important intermediate form. It is, of course, impossible to say with certainty through which of the three-toed genera of the pliocene that lived together the succession came. It is not impossible that the latter species, which appear generically identical, are the descendants of more distinct pliocene types, as the persistent tendency in all the earlier forms was in the same direction. Considering the remarkable development of the group through the tertiary period, and its existence even later, it seems very strange that none of the species should have survived, and that we are indebted for our present horse to the Old World."(7)
PALEONTOLOGY OF EVOLUTION
These and such-like revelations have come to light in our own time--are, indeed, still being disclosed. Needless to say, no index of any sort now attempts to conceal them; yet something has been accomplished towards the same end by the publication of the discoveries in Smithsonian bulletins and in technical memoirs of government surveys. Fortunately, however, the results have been rescued from that partial oblivion by such interpreters as Professors Huxley and Cope, so the unscientific public has been allowed to gain at least an inkling of the wonderful progress of paleontology in our generation.
The writings of Huxley in particular epitomize the record. In 1862 he admitted candidly that the paleontological record as then known, so far as it bears on the doctrine of progressive development, negatives that doctrine. In 1870 he was able to "soften somewhat the Brutus-like severity" of his former verdict, and to a.s.sert that the results of recent researches seem "to leave a clear balance in favor of the doctrine of the evolution of living forms one from another." Six years later, when reviewing the work of Marsh in America and of Gaudry in Pikermi, he declared that, "on the evidence of paleontology, the evolution of many existing forms of animal life from their predecessors is no longer an hypothesis, but an historical fact." In 1881 he a.s.serted that the evidence gathered in the previous decade had been so unequivocal that, had the trans.m.u.tation hypothesis not existed, "the paleontologist would have had to invent it."
Since then the delvers after fossils have piled proof on proof in bewildering profusion. The fossil-beds in the "bad lands" of western America seem inexhaustible. And in the Connecticut River Valley near relatives of the great reptiles which Professor Marsh and others have found in such profusion in the West left their tracks on the mud-flats--since turned to sandstone; and a few skeletons also have been found. The bodies of a race of great reptiles that were the lords of creation of their day have been dissipated to their elements, while the chance indentations of their feet as they raced along the sh.o.r.es, mere footprints on the sands, have been preserved among the most imperishable of the memory-tablets of the world.
Of the other vertebrate fossils that have been found in the eastern portions of America, among the most abundant and interesting are the skeletons of mastodons. Of these one of the largest and most complete is that which was unearthed in the bed of a drained lake near Newburg, New York, in 1845. This specimen was larger than the existing elephants, and had tusks eleven feet in length. It was mounted and described by Dr.
John C. Warren, of Boston, and has been famous for half a century as the "Warren mastodon."
But to the student of racial development as recorded by the fossils all these sporadic finds have but incidental interest as compared with the rich Western fossil-beds to which we have already referred. From records here unearthed, the racial evolution of many mammals has in the past few years been made out in greater or less detail. Professor Cope has traced the ancestry of the camels (which, like the rhinoceroses, hippopotami, and sundry other forms now spoken of as "Old World," seem to have had their origin here) with much completeness.
A lemuroid form of mammal, believed to be of the type from which man has descended, has also been found in these beds. It is thought that the descendants of this creature, and of the other "Old-World" forms above referred to, found their way to Asia, probably, as suggested by Professor Marsh, across a bridge at Bering Strait, to continue their evolution on the other hemisphere, becoming extinct in the land of their nativity. The ape-man fossil found in the tertiary strata of the island of Java in 1891 by the Dutch surgeon Dr. Eugene Dubois, and named Pithecanthropus erectus, may have been a direct descendant of the American tribe of primitive lemurs, though this is only a conjecture.
Not all the strange beasts which have left their remains in our "bad lands" are represented by living descendants. The t.i.tanotheres, or brontotheridae, for example, a gigantic tribe, offshoots of the same stock which produced the horse and rhinoceros, represented the culmination of a line of descent. They developed rapidly in a geological sense, and flourished about the middle of the tertiary period; then, to use Aga.s.siz's phrase," time fought against them." The story of their evolution has been worked out by Professors Leidy, Marsh, Cope, and H.
F. Osborne.
A recent bit of paleontological evidence bearing on the question of the introduction of species is that presented by Dr. J. L. Wortman in connection with the fossil lineage of the edentates. It was suggested by Marsh, in 1877, that these creatures, whose modern representatives are all South American, originated in North America long before the two continents had any land connection. The stages of degeneration by which these animals gradually lost the enamel from their teeth, coming finally to the unique condition of their modern descendants of the sloth tribe, are ill.u.s.trated by strikingly graded specimens now preserved in the American Museum of Natural History, as shown by Dr. Wortman.
All these and a mult.i.tude of other recent observations that cannot be even outlined here tell the same story. With one accord paleontologists of our time regard the question of the introduction of new species as solved. As Professor Marsh has said, "to doubt evolution today is to doubt science; and science is only another name for truth."
Thus the third great battle over the meaning of the fossil records has come to a conclusion. Again there is a truce to controversy, and it may seem to the casual observer that the present stand of the science of fossils is final and impregnable. But does this really mean that a full synopsis of the story of paleontology has been told? Or do we only await the coming of the twentieth-century Lamarck or Darwin, who shall attack the fortified knowledge of to-day with the batteries of a new generalization?
IV. THE ORIGIN AND DEVELOPMENT OF MODERN GEOLOGY
JAMES HUTTON
One might naturally suppose that the science of the earth which lies at man's feet would at least have kept pace with the science of the distant stars. But perhaps the very obviousness of the phenomena delayed the study of the crust of the earth. It is the unattainable that allures and mystifies and enchants the developing mind. The proverbial child spurns its toys and cries for the moon.
So in those closing days of the eighteenth century, when astronomers had gone so far towards explaining the mysteries of the distant portions of the universe, we find a chaos of opinion regarding the structure and formation of the earth. Guesses were not wanting to explain the formation of the world, it is true, but, with one or two exceptions, these are bizarre indeed. One theory supposed the earth to have been at first a solid ma.s.s of ice, which became animated only after a comet had dashed against it. Other theories conceived the original globe as a ma.s.s of water, over which floated vapors containing the solid elements, which in due time were precipitated as a crust upon the waters. In a word, the various schemes supposed the original ma.s.s to have been ice, or water, or a conglomerate of water and solids, according to the random fancies of the theorists; and the final separation into land and water was conceived to have taken place in all the ways which fancy, quite unchecked by any tenable data, could invent.
Whatever important changes in the general character of the surface of the globe were conceived to have taken place since its creation were generally a.s.sociated with the Mosaic: deluge, and the theories which attempted to explain this catastrophe were quite on a par with those which dealt with a remoter period of the earth's history. Some speculators, holding that the interior of the globe is a great abyss of waters, conceived that the crust had dropped into this chasm and had thus been inundated. Others held that the earth had originally revolved on a vertical axis, and that the sudden change to its present position bad caused the catastrophic shifting of its oceans. But perhaps the favorite theory was that which supposed a comet to have wandered near the earth, and in whirling about it to have carried the waters, through gravitation, in a vast tide over the continents.
Thus blindly groped the majority of eighteenth-century philosophers in their attempts to study what we now term geology. Deluded by the old deductive methods, they founded not a science, but the ghost of a science, as immaterial and as unlike anything in nature as any other phantom that could be conjured from the depths of the speculative imagination. And all the while the beckoning earth lay beneath the feet of these visionaries; but their eyes were fixed in air.
At last, however, there came a man who had the penetration to see that the phantom science of geology needed before all else a body corporeal, and who took to himself the task of supplying it. This was Dr.
James Hutton, of Edinburgh, physician, farmer, and manufacturing chemist--patient, enthusiastic, level-headed devotee of science.
Inspired by his love of chemistry to study the character of rocks and soils, Hutton had not gone far before the earth stood revealed to him in a new light. He saw, what generations of predecessors had blindly refused to see, that the face of nature everywhere, instead of being rigid and immutable, is perennially plastic, and year by year is undergoing metamorphic changes. The solidest rocks are day by day disintegrated slowly, but none the less surely, by wind and rain and frost, by mechanical attrition and chemical decomposition, to form the pulverized earth and clay. This soil is being swept away by perennial showers, and carried off to the oceans. The oceans themselves beat on their sh.o.r.es, and eat insidiously into the structure of sands and rocks.
Everywhere, slowly but surely, the surface of the land is being worn away; its substance is being carried to burial in the seas.
Should this denudation continue long enough, thinks Hutton, the entire surface of the continents must be worn away. Should it be continued LONG ENOUGH! And with that thought there flashes on his mind an inspiring conception--the idea that solar time is long, indefinitely long. That seems a simple enough thought--almost a truism--to the twentieth-century mind; but it required genius to conceive it in the eighteenth. Hutton pondered it, grasped its full import, and made it the basis of his hypothesis, his "theory of the earth."
MODERN GEOLOGY
The hypothesis is this--that the observed changes of the surface of the earth, continued through indefinite lapses of time, must result in conveying all the land at last to the sea; in wearing continents away till the oceans overflow them. What then? Why, as the continents wear down, the oceans are filling up. Along their bottoms the detritus of wasted continents is deposited in strata, together with the bodies of marine animals and vegetables. Why might not this debris solidify to form layers of rocks--the basis of new continents? Why not, indeed?
But have we any proof that such formation of rocks in an ocean-bed has, in fact, occurred? To be sure we have. It is furnished by every bed of limestone, every outcropping fragment of fossil-bearing rock, every stratified cliff. How else than through such formation in an ocean-bed came these rocks to be stratified? How else came they to contain the sh.e.l.ls of once living organisms imbedded in their depths? The ancients, finding fossil sh.e.l.ls imbedded in the rocks, explained them as mere freaks of "nature and the stars." Less superst.i.tious generations had repudiated this explanation, but had failed to give a tenable solution of the mystery. To Hutton it is a mystery no longer. To him it seems clear that the basis of the present continents was laid in ancient sea-beds, formed of the detritus of continents yet more ancient.
But two links are still wanting to complete the chain of Hutton's hypothesis. Through what agency has the ooze of the ocean-bed been transformed into solid rock? and through what agency has this rock been lifted above the surface of the water to form new continents? Hutton looks about him for a clew, and soon he finds it. Everywhere about us there are outcropping rocks that are not stratified, but which give evidence to the observant eye of having once been in a molten state.
Different minerals are mixed together; pebbles are scattered through ma.s.ses of rock like plums in a pudding; irregular crevices in otherwise solid ma.s.ses of rock--so-called veinings--are seen to be filled with equally solid granite of a different variety, which can have gotten there in no conceivable way, so Hutton thinks, but by running in while molten, as liquid metal is run into the moulds of the founder. Even the stratified rocks, though they seemingly have not been melted, give evidence in some instances of having been subjected to the action of heat. Marble, for example, is clearly nothing but calcined limestone.
With such evidence before him, Hutton is at no loss to complete his hypothesis. The agency which has solidified the ocean-beds, he says, is subterranean heat. The same agency, acting excessively, has produced volcanic cataclysms, upheaving ocean-beds to form continents. The rugged and uneven surfaces of mountains, the tilted and broken character of stratified rocks everywhere, are the standing witnesses of these gigantic upheavals.
And with this the imagined cycle is complete. The continents, worn away and carried to the sea by the action of the elements, have been made over into rocks again in the ocean-beds, and then raised once more into continents. And this ma.s.sive cycle, In Hutton's scheme, is supposed to have occurred not once only, but over and over again, times without number. In this unique view ours is indeed a world without beginning and without end; its continents have been making and unmaking in endless series since time began.
Hutton formulated his hypothesis while yet a young man, not long after the middle of the century. He first gave it publicity in 1781, in a paper before the Royal Society of Edinburgh:
"A solid body of land could not have answered the purpose of a habitable world," said Hutton, "for a soil is necessary to the growth of plants, and a soil is nothing but the material collected from the destruction of the solid land. Therefore the surface of this land inhabited by man, and covered by plants and animals, is made by nature to decay, in dissolving from that hard and compact state in which it is found; and this soil is necessarily washed away by the continual circulation of the water running from the summits of the mountains towards the general receptacle of that fluid.
"The heights of our land are thus levelled with our sh.o.r.es, our fertile plains are formed from the ruins of the mountains; and those travelling materials are still pursued by the moving water, and propelled along the inclined surface of the earth. These movable materials, delivered into the sea, cannot, for a long continuance, rest upon the sh.o.r.e, for by the agitation of the winds, the tides, and the currents every movable thing is carried farther and farther along the shelving bottom of the sea, towards the unfathomable regions of the ocean.
"If the vegetable soil is thus constantly removed from the surface of the land, and if its place is then to be supplied from the dissolution of the solid earth as here represented, we may perceive an end to this beautiful machine; an end arising from no error in its const.i.tution as a world, but from that destructibility of its land which is so necessary in the system of the globe, in the economy of life and vegetation.
"The immense time necessarily required for the total destruction of the land must not be opposed to that view of future events which is indicated by the surest facts and most approved principles. Time, which measures everything in our idea, and is often deficient to our schemes, is to nature endless and as nothing; it cannot limit that by which alone it has existence; and as the natural course of time, which to us seems infinite, cannot be bounded by any operation that may have an end, the progress of things upon this globe that in the course of nature cannot be limited by time must proceed in a continual succession. We are, therefore, to consider as inevitable the destruction of our land, so far as effected by those operations which are necessary in the purpose of the globe, considered as a habitable world, and so far as we have not examined any other part of the economy of nature, in which other operations and a different intention might appear.
"We have now considered the globe of this earth as a machine, constructed upon chemical as well as mechanical principles, by which its different parts are all adapted, in form, in quality, and quant.i.ty, to a certain end--an end attained with certainty of success, and an end from which we may perceive wisdom in contemplating the means employed.
"But is this world to be considered thus merely as a machine, to last no longer than its parts retain their present position, their proper forms and qualities? Or may it not be also considered as an organized body such as has a const.i.tution, in which the necessary decay of the machine is naturally repaired in the exertion of those productive powers by which it has been formed?
"This is the view in which we are now to examine the globe; to see if there be, in the const.i.tution of the world, a reproductive operation by which a ruined const.i.tution may be again repaired and a duration of stability thus procured to the machine considered as a world containing plants and animals.
"If no such reproductive power, or reforming operation, after due inquiry, is to be found in the const.i.tution of this world, we should have reason to conclude that the system of this earth has either been intentionally made imperfect or has not been the work of infinite power and wisdom."(1)
This, then, was the important question to be answered--the question of the const.i.tution of the globe. To accomplish this, it was necessary, first of all, to examine without prejudice the material already in hand, adding such new discoveries from time to time as might be made, but always applying to the whole unvarying scientific principles and inductive methods of reasoning.
"If we are to take the written history of man for the rule by which we should judge of the time when the species first began," said Hutton, "that period would be but little removed from the present state of things. The Mosaic history places this beginning of man at no great distance; and there has not been found, in natural history, any doc.u.ment by which high antiquity might be attributed to the human race. But this is not the case with regard to the inferior species of animals, particularly those which inhabit the ocean and its sh.o.r.es. We find in natural history monuments which prove that those animals had long existed; and we thus procure a measure for the computation of a period of time extremely remote, though far from being precisely ascertained.
"In examining things present, we have data from which to reason with regard to what has been; and from what actually has been we have data for concluding with regard to that which is to happen hereafter.
Therefore, upon the supposition that the operations of nature are equable and steady, we find, in natural appearances, means for concluding a certain portion of time to have necessarily elapsed in the production of those events of which we see the effects.
"It is thus that, in finding the relics of sea animals of every kind in the solid body of our earth, a natural history of those animals is formed, which includes a certain portion of time; and for the ascertaining this portion of time we must again have recourse to the regular operations of this world. We shall thus arrive at facts which indicate a period to which no other species of chronology is able to remount.
"We find the marks of marine animals in the most solid parts of the earth, consequently those solid parts have been formed after the ocean was inhabited by those animals which are proper to that fluid medium.
If, therefore, we knew the natural history of these solid parts, and could trace the operations of the globe by which they have been formed, we would have some means for computing the time through which those species of animals have continued to live. But how shall we describe a process which n.o.body has seen performed and of which no written history gives any account? This is only to be investigated, first, in examining the nature of those solid bodies the history of which we want to know; and, secondly, in examining the natural operations of the globe, in order to see if there now exist such operations as, from the nature of the solid bodies, appear to have been necessary for their formation.
"There are few beds of marble or limestone in which may not be found some of those objects which indicate the marine object of the ma.s.s. If, for example, in a ma.s.s of marble taken from a quarry upon the top of the Alps or Andes there shall be found one c.o.c.kle-sh.e.l.l or piece of coral, it must be concluded that this bed of stone has been originally formed at the bottom of the sea, as much as another bed which is evidently composed almost altogether of c.o.c.kle-sh.e.l.ls and coral. If one bed of limestone is thus found to have been of marine origin, every concomitant bed of the same kind must be also concluded to have been formed in the same manner.