Taxonomy of the Chipmunks, Eutamias quadrivittatus and Eutamias umbrinus Part 2

_Description._--_Color pattern_: Head Cinnamon mixed with grayish white; stripe on margin of crown Verona-Brown or Bister; ocular stripe Fuscous Black mixed with Sayal Brown; submalar stripe Sayal Brown; ear Fuscous, Sayal Brown along anterior margin and Smoke Gray along posterior margin and on postauricular patch; median dorsal stripe black; lateral dorsal dark stripes Fuscous Black mixed with Russet; outermost dorsal dark stripes slightly darker or indistinguishable from sides in color; dorsal light stripes grayish white with Mikado-Brown along margins; outermost pair of dorsal light stripes nearly pure white; sides Russet mixed with Cinnamon or Ochraceous-Tawny; rump and thighs Smoke Gray mixed with Cinnamon-Buff, with a larger or smaller number of Fuscous Black hairs; antipalmar and antiplantar surfaces of feet Cinnamon-Buff; dorsal surface of tail black; ventral surface of tail Sayal Brown to Tawny; underparts white with dark underfur. _Skull_ and _Baculum_: As in _E. u. umbrinus_.

_Comparisons._--From _E. u. inyoensis_, the subspecies to the west, _E. u. adsitus_ differs in: General tone of upper parts darker; sides darker; interorbital region wider; skull significantly deeper.

For comparison with _E. u. umbrinus_, _E. u. sedulus_, and _E. u. monta.n.u.s_, see the accounts of those subspecies.

_Remarks._--Specimens from West Rim, Zion National Park, 6,500 ft., Washington County, Utah, seem to be intergrades between _E. u.

adsitus_ and _E. u. inyoensis_, and are referable to _E. u. adsitus_.

_Specimens examined._--Total number, 34.

=Utah=: _Beaver Co._: Britts Meadow, Beaver Range Mountains, 8,500 ft., 13 BS. Wayne Co.: Donkey Lake, Boulder Mountain, 10,000 ft., 4 UU. _Garfield Co._: Wildcat R. S., Boulder Mountain, 8,700 ft., 5 UU.

=Arizona=: _Coconino Co._: De Motte Park, Kaibab Plateau, 3 BS; Bright Angel, Kaibab Plateau, 9 BS.

=Eutamias umbrinus sedulus= new subspecies

_Type._--Male, adult, skull, skin, and baculum, No. 158181 U.

S. Nat. Mus. Biol. Surv. Coll.; from Mount Ellen, Henry Mountains, Garfield County, Utah; obtained on October 13, 1908 by W. H. Osgood; original No. 3667.

_Diagnosis._--Size medium; general tone of upper parts dark reddish-brown; ventral surface of tail Ochraceous-Orange; sides Mars Yellow.

_Description._--_Color pattern_: Crown Drab-Gray mixed with Fuscous; upper facial stripe Fuscous Black mixed with Sudan Brown; ocular stripe Sudan Brown mixed with black; submalar stripe Sudan Brown slightly mixed with black; anterior margin of ear Sudan Brown slightly mixed with black; hairs inside pinna, posteriorly, Warm Buff; posterior margin of ear and postauricular patch creamy white; median dorsal dark stripe black with Antique Brown along margins; lateral dorsal dark stripes black mixed with Antique Brown; outermost dorsal dark stripes Xanthine Orange slightly mixed with black; median dorsal light stripes Pale Smoke Gray; outermost dorsal light stripes white slightly mixed with gray; rump and thighs Smoke Gray; sides Mars Yellow; dorsal surface of tail black mixed with Warm Buff; ventral surface of tail Ochraceous-Orange, with black around margin, and Warm Buff around outermost edge; antipalmar and antiplantar surfaces of feet Ochraceous-Buff; underparts creamy white with dark underfur.

_Skull_: Large; braincase moderately inflated; zygomata strong. _Baculum_: As in _E. u. umbrinus_.

_Comparisons._--From _E. u. umbrinus_, the subspecies from the Uinta and northern Wasatch Mountains of Utah, _E. u.

sedulus_ differs in: General tone of upper parts lighter; sides lighter.

From _E. u. adsitus_, the subspecies from the southern Wasatch Range in Utah and Kaibab Plateau in Arizona, _E. u.

sedulus_ differs in: Sides lighter; general tone of upper parts markedly lighter.

From _E. u. inyoensis_, the subspecies from central and northeastern Nevada, and western and northwestern Utah, _E.

u. sedulus_ differs in: Sides lighter (less grayish); general tone of upper parts tawnier.

For comparison with _E. u. monta.n.u.s_, see the account of that subspecies.

_Specimens examined._--Total number, 7 BS, all from the type locality.

=Eutamias umbrinus inyoensis= Merriam

_Eutamias speciosus inyoensis_ Merriam, Proc. Biol.

Soc. Washington, 11:202, 208, July 1, 1897.

_Type._--Male, adult, skull and skin, No. 29387/41462 U. S.

Nat. Mus. Biol. Surv. Coll.; from Black Canyon, 8,200 ft., White Mountains, Inyo County, California; obtained on July 7, 1891, by E. W. Nelson; original No. 1069.

_Diagnosis._--Size medium; sides light; general tone of upper parts light; baculum one of largest in species.

_Description._--_Color pattern_: Head Smoke Gray mixed with Pink-Cinnamon; upper two pairs of facial stripes Fuscous Black or black; submalar stripe Sayal Brown; ear Fuscous or Chaetura-Drab, posterior margin and postauricular patch buffy white; median dorsal dark stripe black with Sayal Brown along margins; lateral dorsal dark stripes black mixed with Sayal Brown or Mikado Brown; outermost dorsal dark stripes Sayal Brown or Mikado Brown mixed with black; sides Ochraceous-Tawny or Tawny; thighs Cinnamon-Buff mixed with Smoke Gray; antipalmar and antiplantar surfaces of feet Cinnamon-Buff; ventral surface of tail Cinnamon-Buff or Ochraceous-Tawny with Fuscous Black around margin and Pinkish Buff around outermost edge; underparts creamy white.

_Skull_: Large; zygomata strong; braincase moderately inflated. _Baculum_: One of largest in species.

_Comparisons._--For comparisons with _E. u. umbrinus_, _E.

u. adsitus_, _E. u. sedulus_, and _E. u. nevadensis_, see the accounts of those subspecies.

_Remarks._--The baculum in _E. u. inyoensis_ is like that in _E.

palmeri_.

_Specimens examined._--Total number, 46.

=Nevada=: _Elko Co._: Head Ackler Creek, N end Ruby Mountains, 1; Steels Creek, N end Ruby Mountains, 1; Summit Secret Pa.s.s, 6,200 ft., Ruby Mountains, 2; Three Lakes, Ruby Mountains, 11; Long Creek, S fork, Ruby Mountains, 4; Harrison Pa.s.s R. S., Green Mountain Canyon, 1; W side Ruby Lake, 6 mi. N Elko Co. line, 3; W side Ruby Lake, 3 mi. N Elko Co. line, 8. _White Pine Co._: Willow Creek, 2 mi. S White Pine Co. line, Ruby Mountains, 6; W side Ruby Lake, 3 mi. S White Pine Co. line, 5; Overland Pa.s.s, E slope Ruby Mountains, 8 mi. S White Pine Co. line, 2.

=Utah=: _Boxelder Co._: Head of George Creek and Clear Creek, 5 mi. S Stanrod, Raft River Mountains, 8,500 ft., 2 UU.

=Eutamias umbrinus nevadensis= Burt

_Eutamias quadrivittatus nevadensis_ Burt, Jour.

Mamm. 12:299, August 24, 1931.

_Type._--Male, adult, skull and skin, No. 15884 Donald R.

d.i.c.key Collection; from Hidden Forest, Sheep Mountains, 8,500 ft., Clark County, Nevada; obtained on July 13, 1929, by W.

H. Burt; original No. 2337.

_Diagnosis._--Size medium; general tone of upper parts grayish; baculum one of the largest of species.

_Description._--"General tone of upperparts grayish; median dorsal stripe, extending from crown between ears to rump, black faintly bordered with 'verona brown'; lateral dark dorsal stripes similar to median stripe, but with anterior one-third deeply suffused with 'verona brown'; central light dorsal stripes grayish, slightly lighter than head and rump; lateral stripes white; head and rump 'pale smoke gray'; postauricular patch grayish white, a narrow margin extending up posterior border of ear; anterior portion of ear 'fuscous black' mixed with 'verona brown' at base and bordered by light gray; ocular stripe black grading into 'verona brown'

in front of ear; submalar stripe nearly obsolete, 'sayal brown'; sides of body grayish washed with 'verona brown'; feet grayish very faintly washed with 'pinkish buff'; dorsal surface of tail black overlaid with 'tilleul buff'; ventral surface of tail 'cinnamon buff' narrowly bordered by black then by 'tilleul buff'; ventral surface of body white." (Burt 1931:299.) Skull similar to that of _E. u. inyoensis_ but differing as indicated below.

_Comparisons._--From _E. u. inyoensis_, the subspecies to the north, _E. u. nevadensis_ differs in: Paler and grayer throughout; tawny areas restricted; gray areas clearer and less suffused; dark facial markings narrower and less distinct; ventral surface of tail distinctly paler; feet lighter, clearer gray; nasals extend farther posteriorly with respect to premaxillae (Burt _loc. cit._).

From _E. u. adsitus_, the subspecies to the northeast, _E. u.

nevadensis_ differs in: Narrower dorsal stripes and facial markings; paler coloration of head, rump, sides, feet, and ventral surface of tail (Burt _op. cit._: 299-300).

_Remarks._--The differences between _E. umbrinus nevadensis_ and _E.

palmeri_, as shown by Burt (_op. cit._) and Hall (1946), are such that one might expect _E. palmeri_ to be a subspecies of _E. umbrinus_.

However, having only the structure of the baculum as evidence additional to that summarized by Hall (_op. cit._), I follow him in according _E. palmeri_ specific status.

_Specimens examined._--None.

=Eutamias umbrinus fremonti= new subspecies

_Type._--Male, adult, skull, skin, and baculum, No. 41790 Univ. Kansas Mus. Nat. Hist.; from 31 mi. N Pinedale, 8,025 ft., Sublette County, Wyoming; obtained on July 8, 1951, by Rollin H. Baker; original No. 1596.

_Diagnosis._--Size large; sides Capucine Yellow; antiplantar surface of hind feet Raw Sienna; postauricular patch grayish white; baculum as in _E. u. umbrinus_.

_Description._--_Color pattern_: Crown Cinnamon-Buff mixed with gray; upper facial stripe Sepia; ocular stripe Chaetura-Drab; submalar stripe Fuscous Black mixed with Sayal Brown; ear black; anterior margin of ear Mars-Yellow, posterior margin grayish white; hairs inside posterior portion of pinna Dresden-Brown; postauricular patch Pale Smoke Gray; median dorsal dark stripe black; lateral dorsal dark stripe black mixed with Sayal Brown; outermost dorsal dark stripe obsolete, Buckhorn-Brown mixed with black; median pair of dorsal light stripes grayish mixed with Buckhorn-Brown; outer pair of dorsal light stripes creamy white; sides Buckhorn-Brown; rump Pale Smoke Gray mixed with Saccardo's Umber; dorsal surface of tail black mixed with Buckhorn-Brown; ventral surface of tail Sayal Brown; outermost edge of tail Light Buff; antipalmar surface of forefeet Warm Buff; antiplantar surface of hind foot Ochraceous-Tawny; underparts creamy white with dark underfur.