Natural History of the Mammalia of India and Ceylon Part 33

NO. 197. LUTRA ELLIOTI.

HABITAT.--Southern Mahratta country.

DESCRIPTION.--The colouring is the same as the last, only a little darker; the distribution of the silvery white is the same; the muzzle is however more depressed than in the last species, and it differs from _L. nair_ by a broader, more arched head, and shorter muzzle.

Dr. Anderson, who distinguishes it by the feature of its skull from the two preceding species, says: "It may be that this otter has a north-westerly distribution, and that it is the species which occurs in the lake at Mount Abu in Rajputana, and also in Sindh and in the Indus."

NO. 198. LUTRA AUROBRUNNEA.

HABITAT.--Nepal.

DESCRIPTION.--Fur of a rich ferruginous brown colour, the upper surface of the head being a deeper brown than the back; the nose is bare; the ears are small and pointed posteriorily. All the strong bristles of the moustache, eyes, cheeks, and chin, are dark brown; claws as in _Lutra_ (_Anderson_). Hodgson says it has a more vermiform body than the rest of Indian otters; tail less than two thirds of the body; nails and toes feebly developed (whence it is cla.s.sed by Gray in the next genus); fur long and rough, rich chestnut-brown above, golden red below and on the extremities.

SIZE.--Head and body, 20 to 22 inches; tail, 12 to 13 inches.

_GENUS AONYX--CLAWLESS OTTERS_.

Muzzle bald, oblong; skull broad, depressed, shorter and more globose than in _Lutra_; the molars larger than in the last genus; flesh tooth larger, and with a large internal lobe; first upper premolar generally absent; feet oblong, elongate; toes slender and tapering; claws rudimentary.

NO. 199. AONYX LEPTONYX.

_The Clawless Otter_ (_Jerdon's No. 102_).

NATIVE NAMES.--_Chusam_, Bhotia; _Suriam_, Lepcha.

HABITAT.--Throughout the Himalayas, also in Lower Bengal and in Burmah.

DESCRIPTION.--"Above earthy brown or chestnut brown; lips, sides of head, chin, throat, and upper part of breast white, tinged with yellowish-grey. In young individuals the white of the lower parts is less distinct, sometimes very pale brownish."--_Jerdon_.

SIZE.--Head and body, 24 Inches; tail, 13.

Mason speaks of this species as common in Burmah, and McMaster mentions his having seen in the Sitang River a colony of white-throated otters smaller than _L. nair_, though larger than _L.

aurobrunnea_, but he did not secure specimens.

AELUROIDEA.

This section includes the Cat family (_Felidae_); the Hyaenas (_Hyaenidae_); two families unknown in India, viz. the _Cryptoproctidae_ and the _Protelidae_; and the Civet family (_Viverridae_).

_FELIDAE--THE CAT FAMILY_.

This family contains the typical carnivores. There is in them combined the greatest power of destruction, accompanied by the simplest mechanism for producing it. All complications of dent.i.tion and digestion disappear. Here are the few scissor-like teeth with the enormous canines, the latter for holding and piercing the life out of their prey, the former for chopping up the flesh into suitable morsels for swallowing. Then the stomach is a simple sac, undivided into compartments, and the intestine is short, not more than three times the length of the body, instead of being some twenty times longer, as in some herbivores. This family has the smallest number of molars, a cla.s.s of tooth which would indeed be useless, for the construction of the feline jaw precludes the possibility of grinding, and therefore a flat-crowned tuberculous tooth would be out of place.

As I have before described it, the jaw of a tiger is incapable of lateral motion. The condyle of the lower jaw is so broad, and fits so accurately into its socket, the glenoid cavity, that there can be no departure from the up and down scissor-like action. The true Cats have, therefore, only one molar on each side of each jaw; those in the upper jaw being merely rudimentary, and placed almost at right angles to the rest of the teeth, and seem apparently of little use; those of the lower jaw are large and trenchant, cutting against the edge of the third upper premolar.

[Ill.u.s.tration: Skull of Tiger (side view).]

It may interest my readers to know which are premolars and which are molars. This can be decided only by dissection of the jaw of a young animal. True molars only appear as the animal approaches the adult stage. They are never shed, as are all the rest of the teeth, commonly called milk teeth. The deciduous or milk teeth are the incisors, canines, and premolars; they drop out and are replaced, and behind the last premolar comes up the permanent molar.

Another peculiar feature of the Cat family is the power of sheathing their talons. Claws to a cat are of as great importance to him in the securing of his prey as are his teeth. The badger is a digger, Hodge, who carries his mattock on his shoulder; but the feline is the free-lance whose sword must be kept keen in its scabbard, so by a peculiar arrangement of muscles the points of the claws are kept off the ground, while the animal treads noiselessly on soft pads.

Otherwise by constant abrasion they would get so blunted as to fail in their penetrating and seizing power. I give here an ill.u.s.tration of the mechanism of the feline claw. In the upper sketch the claw is retracted or sheathed; in the lower it is protruded as in the act of striking.

[Ill.u.s.tration: Tendons of Tiger's toe.]

[Ill.u.s.tration: Tiger's auditory apparatus.]

The senses of hearing and smell are much developed, and the bulb of the ear (_bulla tympani_) is here found of the largest dimensions.

I have once before alluded to this in writing of the bears, in whom this arrangement is deficient. I give here a section of the auditory apparatus. I do not know whether the engraver has effectually rendered my attempt at conveying an idea, based as it is on dissections by Professor Flower; but if he has failed I think the fault lies in the shakiness of my hand in attempting the fine shading after nearly breaking a saw and losing my temper over a very tough old skull which I divided before commencing my ill.u.s.tration. The great cavity is the _bulla tympani_ or bulb of the ear; _a m_ is the _auditory meatus_ or external hole of the ear. On looking into a dry skull the pa.s.sage seems to be of no great depth, nor can an instrument be pa.s.sed directly from the outside into the great tympanic cavity, the hindrance being a wall of bone, _s_, the _septum_ which divides the _bulla_ into two distinct chambers, the reason for which is not very clear, except that one may suppose it to be in some measure for acoustic purposes, as all animals with this development are quick of hearing. The communication between the two chambers lies in a narrow slit over the _septum_, the Eustachian tube, _e_, being on the outside of the _septum_ and between it and the tympanum or ear drum, _t_.

The above are the chief characteristics of the family. For the rest we may notice that they have but a rudimentary clavicle imbedded among the muscles; the limbs are comparatively short, but immensely muscular; the body lithe and active; the foot-fall noiseless; the tongue armed with rough papillae, which enables them to rasp the flesh off bones, and their vision is adapted for both night and day.

None of them are gregarious, as in the case of dogs and wolves. One hears sometimes of a limited number of lions and tigers being seen together, but in most cases they belong to one family, of which the junior members have not been "turned off on their own hook" as yet.

_GENUS FELIS_.

NO. 200. FELIS LEO.

_The Lion_ (_Jerdon's No. 103_).

NATIVE NAMES.--_Sher-babbar_, _Singh_, _Unthia-bagh_.

HABITAT.--Guzerat and Central India.

[Ill.u.s.tration: _Felis leo_ (Indian variety).]

DESCRIPTION.--The lion is almost too well known to need description, and there is little difference between the Asiatic and African animal.

It may, however, be generally described as being distinguished from other Cats by its uniform tawny colour, flatter skull, which gives it a more dog-like appearance, the s.h.a.ggy mane of the male, and by the tufted tail of both s.e.xes.

SIZE.--From nose to insertion of tail, 6 to 6-1/2 feet; tail, 2-1/2 to 3 feet; height, 3-1/2 feet.

The weight of one measured by Captain Smee, 8 feet 9-1/2 inches, was (excluding the entrails) thirty-five stone. This must be the one alluded to by Jerdon, but he does not state the extraction of the viscera, which would add somewhat to the weight.

Young lions when born are invariably spotted; and Professor Parker states that there were in the Zoological Gardens in 1877 three lions which were born in the menagerie about ten years previously, and which showed "indistinct, though perfectly evident, spots of a slightly darker tawny than the general ground-tint on the belly and flanks." He adds: "This is also the case with the puma, and it looks very much as if all the great Cats were descended from a spotted ancestor." The more dog-like head of the lion is well known to all who have studied the physiognomy of the Cats, and I have not only noticed it in drawing the animal, but have seen it alluded to in the writings of others. It was not, however, till lately that I had an opportunity of comparing the skulls of the lion and tiger in the Calcutta Museum, and I am indebted to Mr. c.o.c.kburn of the museum, not only for the trouble he took in getting out the various skulls, but for his a.s.sistance in pointing out certain peculiarities known to him, but of which I was at the time ignorant. That the skull of the lion is flatter than, and wants the bold curve of, those of the tiger, leopard and jaguar, is a well-known fact, but what Mr.

c.o.c.kburn pointed out to me was the difference in the maxillary and nasal sutures of the face. A glance at two skulls placed side by side would show at once what I mean. It would be seen that the nasal bones of the tiger run up higher than those of the lion, the apices of whose nasal and maxillary sutures are on a level. On leaving the museum I compared the tiger skulls in my possession with accurate anatomical drawings which I have of the osteology of the lion, and the result was the same. It is said that there is also a difference in the infra-orbital foramen of the two animals, but this I have failed to detect as yet, though a.s.serted by De Blainville in his magnificent work on osteology ('Osteographie').

From all that has been written of the African and Indian lions I should say that the tiger was the more formidable of the two, as he is, I believe, superior in size. About twenty-two years ago my attention was drawn to this subject by the perusal of Mr. Blyth's article on the _Felidae_ in the old _India Sporting Review_ of 1856-57. If I am not mistaken there was at that time (1861) a fine skeleton of a lion in the museum, as well as those of several tigers, which I measured. I had afterwards opportunities of observing and comparing skeletons of the two animals in various museums in Europe, though not in my own country, for my stay in England on each occasion of furlough was brief, and in almost every instance I found the tiger the larger of the two. The book in which I recorded my observations, and which also contained a number of microscopic drawings of marine infusoria, collected during a five months' voyage, was afterwards lost, so I cannot now refer to my notes.

I believe there was once a case of a fair fight between a well-matched lion and tiger in a menagerie (Edmonds's, I think). The two, by the breaking of a part.i.tion, got together, and could not be separated.

The duel resulted in the victory of the tiger, who killed his opponent.

The lion seems to be dying out in India, and it is now probably confined only to Guzerat and Cutch. I have not been an attentive reader of sporting magazines of late years, and therefore I cannot call to mind any recent accounts of lion-killing in India, if any such have been recorded. At the commencement of this century lions were to be found in the North-West and in Central India, including the tract of country now termed the Central Provinces. In 1847 or 1848 a lioness was killed by a native shikari in the Dumoh district.

Dr. Spry, in his 'Modern India,' states that, when at Saugor in the Central Provinces in 1837, the skin of a full-grown male lion was brought to him, which had been shot by natives in the neighbourhood.

He also mentions another lioness shot at Rhylee in the Dumoh district in 1834, of which he saw the skin. Jerdon says that tolerably authentic intelligence was received of the presence of lions near Saugor in 1856; and whilst at Seonee, within the years 1857 to 1864, I frequently heard the native shikaris speak of having seen a tiger _without stripes_, which may have been of the present species. The indistinct spots on the lion's skin (especially of young lions), to which I have before alluded, were noticed in the skin of the lioness shot at Dumoh in 1847. The writer says: "when you place it in the sun and look sideways at it, some very faint spots (the size of a shilling or so) are to be seen along the belly."

Lions pair off at each season, and for the time they are together they show great attachment to each other, but the male has to fight for his spouse, who bestows herself on the victor. They then live together till the young are able to shift for themselves. The lioness goes with young about fifteen or sixteen weeks, and produces from two to six at a litter. But there is great mortality among young lions, especially about the time when they are developing their canine teeth.