Geographic Distribution and Taxonomy of the Chipmunks of Wyoming Part 4

Mean (9) 31.5 17.6 15.2 9.7 193 85.6 4.78 16.52 Min. [Male] 30.3 17.0 14.9 8.9 183 77.0 4.58 15.63 Max. 32.4 18.2 15.5 10.6 203 91.0 5.12 17.37 Mean (8) 32.2 18.0 15.4 9.7 203 85.7 4.86 16.50 Min. [Female] 31.1 17.6 15.0 9.2 194 79.0 4.64 15.44 Max. 33.4 18.5 15.8 10.2 212 92.0 5.11 17.21 --------------+------+------+------+------+-----+-------+------+-------+ _E. a. luteiventris_, near Moran, Teton Co.

Mean (20) 33.6 18.2 15.5 10.6 212 94.8 5.14 17.27 Min. [Male] 32.2 17.4 14.9 9.8 198 87.0 4.86 16.42 Max. 35.2 18.7 16.2 12.1 221 108.0 5.37 18.39 Mean (10) 33.8 18.5 15.5 11.1 217 91.7 5.13 17.47 Min. [Female] 33.4 18.1 15.1 10.5 203 81.0 5.06 16.89 Max. 34.7 19.0 16.0 11.5 225 100.0 5.32 18.33 --------------+------+------+------+------+-----+-------+------+-------+ _E. d. utahensis_, W side Green River, 1 mi. N Utah border.

Mean (4) 34.7 18.9 16.4 10.8 197 84.5 5.08 17.91 Min. [Male] 34.7 18.7 16.4 10.5 191 81.0 5.00 17.77 Max. 34.8 19.2 16.4 11.1 203 88.0 5.15 18.06 Mean (2) 36.0 19.5 16.3 11.3 211 88.0 5.25 18.87 Min. [Female] 35.5 19.4 16.2 11.3 210 86.0 5.22 18.73 Max. 36.6 19.7 16.4 11.4 212 90.0 5.28 19.02 --------------+------+------+------+------+-----+-------+------+-------+ _E. u. umbrinus_, Mts. S Robertson, Uinta Co.

Mean (11) 34.7 18.9 15.7 10.9 218 96.2 5.13 18.04 Min. [Male] 34.3 18.3 15.6 10.3 215 81.0 4.79 17.57 Max. 35.2 19.4 16.0 11.7 228 112.0 5.42 18.59 Mean (4) 35.1 19.2 15.9 11.0 224 96.4 5.17 18.46 Min. [Female] 34.9 19.2 15.7 10.3 204 90.0 5.11 18.31 Max. 35.4 20.0 16.2 11.8 234 100.0 5.22 18.98 --------------+------+------+------+------+-----+-------+------+-------+ _E. u. fremonti_, Togwotee Pa.s.s, Fremont Co.

Mean (8) 35.6 19.3 15.9 11.4 223 99.0 5.34 19.17 Min. [Male] 35.2 18.9 15.8 11.1 216 95.0 5.22 18.72 Max. 36.5 19.7 16.1 11.8 243 111.0 5.57 19.78 Mean (6) 35.3 19.6 15.9 11.3 229 101.0 5.40 19.02 Min. [Female] 34.5 19.3 15.7 10.9 223 92.0 5.35 18.37 Max. 36.0 20.0 16.5 12.0 239 110.0 5.44 19.51 --------------+------+------+------+------+-----+-------+------+-------+ _E. u. monta.n.u.s_, near Longs Peak, Boulder Co., Colorado.

Mean (5) 35.2 18.8 15.5 10.8 226 96.0 5.20 18.29 Min. [Male] 34.7 18.4 15.2 10.1 215 93.0 5.03 17.80 Max. 36.8 19.4 16.2 11.5 232 115.0 5.53 19.36 Mean (6) 35.7 19.1 15.6 10.9 226 98.0 5.28 18.67 Min. [Female] 35.1 18.8 15.1 10.3 215 89.0 5.06 18.09 Max. 36.5 19.5 16.0 11.5 231 105.0 5.58 19.35 --------------+------+------+------+------+-----+-------+------+-------+

Eutamias amoenus (J. A. Allen)

_Diagnosis._--Size medium; over-all tone of upper parts often grayish olive; baculum small or medium, slender; tip of baculum 30 to 38 per cent of length of shaft; skull medium, narrow across zygomata.

_Comparisons._--From _E. dorsalis utahensis_, the only subspecies of this species in Wyoming, _E. amoenus luteiventris_ differs in: Dorsal light and dark stripes distinct; over-all tone of upper parts less grayish (more tawny); tip of baculum less than 38 per cent of length of shaft in adult specimens.

From _E. umbrinus fremonti_, the only subspecies of this species which occurs in the same area with _E. amoenus_ in Wyoming, _E. a.

luteiventris_ differs in: Smaller size; tawny underparts; base of baculum not noticeably widened.

For comparisons with _E. minimus_ see the account of that species.

Eutamias amoenus luteiventris (J. A. Allen)

_Tamias quadrivittatus luteiventris_ J. A. Allen, Bull. Amer. Mus.

Nat. Hist., 3:101, June, 1890.

_Eutamias amoenus luteiventris_, Howell, Jour. Mamm., 3:183, August 4, 1922.

_Type._--Male, adult, skull and skin, No. 11991/37996 (NM); from "Chief Mountain Lake" [Waterton Lake], 3-1/2 mi. N United States-Canadian Boundary, Alberta; obtained on August 24, 1874, by Elliot Coues; original No. 4596.

_Diagnosis._--General tone of upper parts ochraceous; underparts strongly buffy; tip of baculum in adult specimens, more than 30 per cent and less than 38 per cent of length of shaft.

_Description._--_Color pattern_: Crown Cinnamon mixed with Smoke Gray; upper two facial stripes black; submalar stripe Fuscous or Fuscous Black mixed with Ochraceous Tawny; anterior margin of ear Ochraceous Tawny; posterior margin of ear and postauricular patch Light Buff or buffy white; hairs inside posterior part of pinna Ochraceous Tawny; median dorsal dark stripe black; lateral pair of dorsal dark stripes black and mixed with Tawny, frequently brownish; median pair of dorsal light stripes white tinged with Pale Smoke Gray; lateral pair of dorsal light stripes creamy white; sides Tawny or Ochraceous Tawny; rump and thighs Dark Smoke Gray strongly mixed with Cinnamon Buff; dorsal surface of tail Fuscous Black mixed with Clay Color; ventral surface of tail Light Ochraceous Tawny, with Fuscous Black around margin and Clay Color around outermost edge; antipalmar and antiplantar surfaces of feet Cinnamon or Cinnamon Buff; underparts Cinnamon Buff or Light Ochraceous Buff. _Skull_: Size medium; moderately narrowed across zygomata. _Baculum_: Slender; not noticeably broadened at base; tip more than 30 per cent of length of shaft.

_Remarks._--Although there are no records of this subspecies from the Wind River Mountains, it probably occurs there.

The niche that this subspecies occupies is similar to that of _E. m.

consobrinus_ as shown by the fact that these two subspecies have been taken together at the same places.

Specimens of _E. a. luteiventris_ and _E. umbrinus fremonti_ have been taken together at the same places.

In general, _E. m. consobrinus_ occurs in open country and at the edges of forests, whereas _E. u. fremonti_ occurs in the forest. _E. a.

luteiventris_ occurs in the intermediate habitat, that is to say, not far into the forest, and not so far out into the open as _E. m.

consobrinus_.

_Specimens examined._--Total number, 83.

_Yellowstone Park_: Unspecified, 2.

_Park Co._: 31-1/2 mi. N and 36 mi. W Cody, 6,900 ft., 6; 29 mi. N and 31 mi. W Cody, 7,200 ft., 1; 28 mi. N and 30 mi. W Cody, 7,200 ft., 1; 16-1/4 mi. N and 17 mi. W Cody, 5,625 ft., 3; 25 mi. S and 28 mi. W Cody, 6,350 ft., 2.

_Teton Co._: Two Ocean Lake, 2 (1 FC); Whetstone Creek, 8 (MM); Emma Matilda Lake, 1 (FC); Pacific Creek Road, 2-1/2 mi. E Moran, 1 (FC); Two Ocean Lake Road, 2 (FC); 2 mi. E Moran, 1 (FC); 2-1/2 mi.

E and 1/4 mi. N Moran, 6,230 ft., 8; Pacific Creek, 1 (MM); junction of Two Ocean Lake Road and U.S. Highway 187, 2 (FC); Signal Mountain Road, 1 (FC); Leigh Lake, 9 (MM); Indian Paint Brush Canyon, Teton Park, 1 (MM); Teton National Park, 3; 3 mi. E and 1/4 mi. S Moran, 6,200 ft., 1; 3-3/4 mi. E and 1 mi. S Moran, 6,200 ft., 8; 2-1/2 mi. N and 3-1/2 mi. E Moran, 7,225 ft., 1; Timbered Island, 6,750 ft., 4 mi. N Moose, 5; Bar BC Ranch, 6,500 ft., 2-1/2 mi. NE Moose, 9; Grand Teton, 9,000 ft., Teton Park, 1 (MM); Upper Arizona Creek, Jackson, 1 (MM).

_Lincoln Co._: 3 mi. N and 11 mi. E Alpine, 5,650 ft., 2.

_Additional records_ (Howell 1929:69): _Yellowstone Park_: Mammoth Hot Springs; Roaring Mountain; Bunsen Peak; Yancey; Apollinaris Spring; Canyon; Yellowstone Lake; Upper Geyser Basin; Old Faithful.

_Park Co._: Near head of Clarks Fork; Pahaska, N Fork Shoshone River at Grinnell Creek; Valley. _Teton Co._: Moran; Teton Mountains; Teton Pa.s.s. _Lincoln Co._: Afton, Salt River Mountains; head of La Barge Creek, 9,100 ft.; Salt River Mountains, 10 mi. SE Afton. _Sublette Co._: Merna; Stanley.

Eutamias dorsalis (Baird)

_Diagnosis._--Size medium to large; general tone of upper parts Smoke Gray; dorsal stripes indistinct or obsolete; often brightly colored at base of tail; keel of baculum proportionally high, approximately 1/3 of length of tip; skull longer than 34.5 mm.

_Comparisons._--From _E. umbrinus_, _E. dorsalis_ differs in: Dorsal stripes faint; skull smaller; base of baculum not noticeably expanded; general tone of upper parts grayer.

For comparisons with _E. minimus_ and _E. amoenus_ see the accounts of those species.

[Ill.u.s.tration: FIG. 2. Known occurrences and probable geographic distribution of _Eutamias amoenus_ and _Eutamias dorsalis_ in Wyoming. See figure 1 for explanation of symbols.

1. _E. amoenus luteiventris_ 2. _E. dorsalis utahensis_]

Eutamias dorsalis utahensis Merriam

_Eutamias dorsalis utahensis_ Merriam, Proc. Biol. Soc. Washington, 11:210, July 1, 1897.

_Type._--Male, adult, skull and skin, No. 186457 (NM); from Ogden, Weber County, Utah; obtained on October 9, 1888, by Vernon Bailey; original No. 289.

_Diagnosis._--Size medium; dorsal stripes faint; baculum not noticeably widened at base.

_Description._--_Color pattern_: Crown Pale Smoke Gray mixed with Cinnamon; upper facial stripe Fuscous; other facial stripes Sayal Brown mixed with Fuscous or Fuscous Black; anterior margin of ear Ochraceous Tawny; posterior margin of ear and postauricular patch grayish white; median dorsal dark stripe Fuscous or black; other dorsal dark stripes black and mixed with gray, sometimes barely discernible; dorsal pair light stripes Smoke Gray; lateral pair of light stripes creamy white; rump and thighs Pale Smoke Gray mixed with Cinnamon; dorsal surface of tail Fuscous Black mixed with Tilleul Buff; underside of tail Cinnamon Buff or Pinkish Buff, Fuscous Black around margin and Tilleul Buff around outermost edge; antipalmar and antiplantar surfaces of feet Cinnamon Buff; underparts creamy white; sides Pinkish Cinnamon or Light Pinkish Cinnamon. _Skull_: Size medium; braincase well inflated; zygomata strong, moderately appressed to cranium. _Baculum_: Small; keel approximately 1/3 of length of tip.

_Remarks._--Only a few specimens of this subspecies have ever been taken in Wyoming. Little is known about the habits of this chipmunk, which normally is shy and wary.

_Specimens examined._--Total number, 6.

_Sweet.w.a.ter Co._: W side Green River, 1 mi. N Utah border, 6.

_Additional records_ (Howell 1929:134): _Sweet.w.a.ter Co._: Green River, 4 mi. NE Linwood, Utah.

[Ill.u.s.tration: FIG. 3. Known occurrences and probable geographic distribution of the subspecies of _Eutamias umbrinus_ in Wyoming. See figure 1 for explanation of symbols.

1. _E. u. umbrinus_ 2. _E. u. fremonti_ 3. _E. u. monta.n.u.s_]